应用世代分析法估算鱼类自然死亡率的初步研究A Preliminary Study on the Estimation of Natural Mortality Using Cohort Analysis Method
王迎宾,刘群
摘要(Abstract):
根据已开发鱼类种群的自然死亡率(M)在鱼类资源评估中的重要作用,讨论了1种应用实际种群分析(VPA)(亦即世代分析方法)估算鱼类自然死亡率的方法。文中假设了捕捞努力量(E)已知和未知2种情况,并同时求出了其他种群参数,如可捕系数,补充量等。在蒙特卡罗模拟分析中,假设了4种模拟渔业(管理良好的渔业,掠夺式渔业,恢复性渔业和稳定性渔业)对该方法进行了验证。并将von-Bertalanffy生长方程(VBGF)引入VPA模型来估算鱼类的自然死亡率。当C V等于20%时,在4种模拟渔业中鱼类各龄自然死亡率(Ma)的EP均小于10%。在已知单位捕捞努力量渔获量数据时,计算北大西洋长鳍金枪鱼(Thunnus alalunga)的M等于0.119,这个结果比先前假定的值(0.3)小,但考虑到该鱼种所承受的高的捕捞压力作者认为该估计值是合理的。
关键词(KeyWords): 自然死亡率;实际种群分析;模拟分析;长鳍金枪鱼
基金项目(Foundation): 国家自然科学基金项目(30271025)资助
作者(Author): 王迎宾,刘群
DOI: 10.16441/j.cnki.hdxb.2005.05.006
参考文献(References):
- [1]Tanasichuk R W.Age-specific natural mortality rates of adult Pacificherring(Clupea pallasi)from southern British Columbia[J].Can JFish Aquat Sci,2000,57:2258-2266.
- [2]Zheng J.Uncertainties of natural mortality estimates for easternBering Sea snow crab,Chionoecetes opilio[J].Fish Res,2003,65:411-425.
- [3]QuinnⅡT J,Deriso R B.Quantitative Fish Dynamics[M].NewYork:Oxford University Press,1999.
- [4]Gulland J A.Estimation of mortality rates.Annex to Rep.Arc.Fish.Working Group[R].Copenhagen:Int.Counc.Expolr.SeaC.M.1965.9.
- [5]ICES.Report of the workshop on methods of fish stock assessment[R].Reykjavik:ICES C M 1988/Assess,1988.117.
- [6]ICES.Report of the workshop on methods of fish stock assessment[R].St John’s:ICES C M 1991/Assess,1991.149.
- [7]ICES.Report of the workshop on methods of fish stock assessment[R].Copenhagen:ICES C M 1993/Assess,1993.86.
- [8]Lassen H.Estimation of stock size and fishing mortality when data oncatches and on stock abundance are available[R].Copenhagen:IntCounc Explor Sea Stat Comm,CM,Doc.1983.14.
- [9]Deriso R B,QuinnⅡT J,Neal P R.Catch-age analysis with auxil-iary information[J].Can J Fish Aquat Sci,1985,42:815-824.
- [10]Shepherd J G.Extended survivors analysis:An improved method forthe analysis of catch-at age data abundance indices[J].ICES Journalof Marine Science,1999,56:584-591.
- [11]Jennings S,Kaiser M J,Reynolds J D.Marine Fisheries Ecology[M].Malden:USA Blackwell Science Ltd.,2001.417.
- [12]Gavaris S.An Adaptive Framework for the Estimation of PopulationSize[C].Canada:Canadian Atlantic Fisheries Sci.Adv.Comm.Res.Doc.,1988.
- [13]Kimura D K.Variability,tuning,and simulation for the Doubleday-Deriso catch-at-age model[J].Can J Fish Aquat Sci,1989,46:941-949.
- [14]Liu Q,Pitcher T J.Application of a constrained optimization methodto the statistical analysis of fisheries catch-at-age data[J].Fish Res,1995,22:29-41.
- [15]Myers R A,QuinnⅡT J.Estimating and testing non-additivity infishing mortality:implications for detecting a fisheries collapse[J].Can J Fish Aquat Sci,2002,59:597-601.
- [16]Ricker W E.Computation and Interpretation of Biological Statisticsof Fish Populations[s.l.]:Bull Fish Res Board Can,1975.
- [17]Pope J G,Shepherd J G.A simple method for the consistent inter-pretation of catch at age data[J].J Cons Int Explor Mer,1982,40:129-151.
- [18]Sparre P,Venema S C.Introduction to Tropical Fish Stock Assess-ment-Part 1:Manual[R].Rome:FAO-Food and Agriculture Orga-nization of the United Nations.In,1998.407.
- [19]Beyer J E,Kirchner C H,Holtzhausen J A.A method to determinesize-specific natural mortality applied to westcoast steenbras(Lithog-nathus aureti)in Namibia[J].Fish Res,1999,41:133-153.
- [20]Beyer J E.Recruitment stability and survival-simple size-specific the-ory with examples from the early life dynamics of marine fish[J].Dana,1989,7,45-147.
- [21]詹秉义.渔业资源评估[M].北京:中国农业出版社,1995.353.
- [22]Press W H,Flannery B P,Teukolsky S A,et al.Numerical Recipes[M].Cambridge,U.K.:Cambridge University Press,1986.
- [23]ICCAT.Report of Albacore Stock Assessment Session[R].Madrid:SCRS/2003/017.Assess:2003.15-20.
- [24]Shepherd J G,Pope J G.Dynamic pool models 1:Interpreting thepast using virtual population analysis[A].Qaul J B Hart,John DReynolds.Hand Book of Fish Biology and Fisheries:Volume 2 Fish-eries[M].USA:Blackwell Science Ltd.Malden,2002.410.
- [25]Fu C H,QuinnⅡT J.Estimability of natural mortality and otherpopulation parameters in a length-based model:Pandalus borealisinKachemak Bay,Alaska[J].Can J Fish Aquat Sci,2000,57:2420-2432.
- [26]Schnute J T,Richards L J.The influence of error on population esti-mates from catch-age models[J].Can J Fish Aquat Sci,1995,52:2063-2077.
- [27]ICCAT.Report for Biennial Period 1998-99[R].Madrid:SCRS/1999/I Assess,1999.289.
- [28]ICCAT.Report for Biennial Period 1998-99[R].Madrid:SCRS/2000/ⅡAssess:2000.175.
- [29]FAO.Review of the State of World Fishery Resources:MarineFisheries[R].Rome:FAO.2003.103.
- [30]Doubleday W G.A least squares approach to analyzing catch-at-agedata[J].Res Bull Int NW Atl Fish,1976,12:69-81.
- [31]倪建峰,刘群.剩余产量模型在不同渔业中的应用[J].海洋湖沼通报,2004,1:60-67.
- [32]Paloheimo J E.Estimation of mortality rates in fish populations[J].Trans Am Fish,Soc,1980,1094:378-386.
- [33]王迎宾,刘群.鱼类自然死亡率的估算及其影响因子的探讨[J].中国海洋大学学报,2005,35(1):20-24.
- [34]Ricker W E.Effects of size-selective mortality and sampling bias onestimates of growth,mortality,production,and yield[J].J FishRes Board Can,1969,26:479-541.
- [35]Ware D M.Relation between egg size,growth and natural mortalityof larval fish[J].J Fish Bull,1975,86(1):25-43.
- [36]Peterson I,Wroblewski S J.Mortality rate of fishes in the pelagic e-cosystem[J].Can J Fish Aquat Sci,1984,41:1117-1120.
- [37]David A F,John J H,John R S.MULTIFAN-CL:a length-based,age-structured model for fisheries stock assessment,with applicationto South Pacific albacore,Thunnus alalunga[J].Can J Fish AquatSci,1998,55:2105-2116.
- [38]Torres F S B.Jr.Tabular data on marine fishes from Sourth Africa,PartⅡ:Growth parameters[J].Fishbyte,1991,9(2):37-38.
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